Guangdong Key Laboratory for Genome Stability & Disease Prevention and Carson International Cancer Center, Marshall Laboratory of Biomedical Engineering, Shenzhen University Medical School, Shenzhen University, Shenzhen, Guangdong, China.

Shenzhen Key Laboratory for Systemic Aging and Intervention(SKL-SAI), School of Basic Medical Sciences, Shenzhen University Medical School, Shenzhen, Guangdong, China.

Department of Immunology, University of Tübingen, Tübingen, Germany.

Departments of Biology and Medicine, University of Rochester, Rochester, NY, USA.

University of Chinese Academy of Sciences, Beijing, China.

Department of Epigenetics, Van Andel Institute, Grand Rapids, MI, USA.

Department of Medicine, Center for Advanced Gerotherapeutics, Cedars-Sinai Medical Center, Los Angeles, CA, USA.

School of Biomedical Sciences, University of Hong Kong, Hong Kong, China.

Institute for Research on Cancer and Aging of Nice (IRCAN), INSERM, Université Côte d'Azur, Centre National de la Recherche Scientifique(CNRS), Nice, France.

Life Sciences Institute, Zhejiang University, Hangzhou, Zhejiang, China.

Department of Oncology, Mayo Clinic, Rochester, MA, USA.

Tongji University School of Medicine, Shanghai, China.

Department of Medicine, Breast Unit, The Royal Marsden NHS Foundation Trust, Sutton, UK.

Cellular senescence, Cancer & Aging Lab, Cancer Research Center of Lyon, Lyon, France.

Department of Microbiology Immunology and Parasitology, UNIFESP, Federal University of São Paulo, São Paulo, Brazil.

Shanghai Key Laboratory for Tumor Microenvironment and Inflammation, Department of Biochemistry and Molecular Cell Biology, Shanghai Jiao Tong University School of Medicine, Shanghai, China. Institute of Aging & Tissue Regeneration, Ren-Ji Hospital, Shanghai Jiao Tong University School of Medicine, Shanghai, China. Hainan Academy of Medical Sciences, Haikou, Hainan, China.

Gustave Roussy, Université Paris-Saclay, Villejuif, France.

IFOM, The AIRC Institute of Molecular Oncology, Milan, Italy.

National Clinical Research Center for Geriatrics, State Key Laboratory of Biotherapy, West China Hospital, Sichuan University, Chengdu, Sichuan, China.

Department of Oncology, MZB Foundation for Cancer Research, New York, NY, USA.

Geriatric Unit, Lyon Sud Hospital, Pierre-Benite, France.

Department of Medical Oncology, Local Health Unit of Western Lisbon, Lisbon, Portugal.

Université Paris Cité, INSERM U970, PARCC, Paris, France.

Department of Virology-Oncology, Apex Institute of Medical Research, Besançon, France.

Cordeliers Research Center, Université de Paris Cité, Paris, France.

Department of Medicine, Faculty of Medicine and Health Sciences, University of Sherbrooke, Sherbrooke, QC, Canada.

The Second Affiliated Hospital of Guangzhou University of Chinese Medicine, Guangzhou, Guangdong Province, China.

The First Affiliated Hospital, School of Medicine, Zhejiang University, Hangzhou, Zhejiang, China.

Department of Urology, Johns Hopkins University, Baltimore, MD, USA.

Department of Immunology, Leiden University Medical Center, Oegstgeest, The Netherlands.

Center for Genomic Integrity, Institute for Basic Science, Ulsan National Institute of Science and Technology, Ulsan, South Korea.

Department of Hematology and Stem Cell Transplantation, University Hospital Essen, University Duisburg-Essen, Hufelandstrasse, Essen, Germany.

Department of Medical Oncology, Leiden University Medical Center, Leiden, The Netherlands.

Department of Immunology, Universidad De Cordoba, Cordoba, Spain.

Department of Physiology, Immunology Unit, University of Extremadura, Cáceres, Spain.

Department of Biochemistry, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Department of hematology, oncology and immunotherapy cantonal hospital Baselland, Medical University Clinics, Basel, Switzerland.

Department of Biomedical Engineering, Faculty of Engineering, Hong Kong Polytechnic University, Hong Kong, China.

School of Life Sciences, Sichuan University, Chengdu, Sichuan, China.

Westlake Laboratory of Life Sciences and Biomedicine, Hangzhou, Zhejiang, China.

Suzhou Institute of Systems Medicine, Chinese Academy of Medical Sciences & Peking Union Medical College, Suzhou, Jiangsu, China.

China National Center for Bioinformation, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China.

Institute of Aging Research, School of Medicine, Hangzhou Normal University, Hangzhou, Zhejiang, China.